Humboldt State University

William “Tim” Bean, Ph.D.

707 826 3658


  • Bean, W.T., D. Tange*, and S. Osborn. 2016. A suitability model for white-footed voles with insights into habitat associations at the southern boundary of their range. Northwestern Naturalist 97:105-112.
  • Dulava, S., W.T. Bean, and O. Richmond. 2015. Applications of unmanned aircraft systems (UAS) for waterbird surveys. Environmental Practice 17:201-210.
  • Bean, W.T., H.S. Butterfield, R. Stafford, and J.S. Brashares. 2014. Multi-step realized distribution models suggest bottom-up control of an endangered rodent. PLoS ONE 9:e106638
  • Bean, W.T., L.P. Prugh, R. Stafford, H.S. Butterfield, M. Westphal, and J.S. Brashares. 2014. Species distribution models of an endangered rodent offer incomplete measures of habitat quality at multiple scales. Journal of Applied Ecology 51:1116-1125.
  • Bean, W.T., R. Stafford, H.S. Butterfield, L.R. Prugh and J.S. Brashares. 2012. An evaluation of monitoring methods for giant kangaroo rats. Wildlife Society Bulletin 36:587-593.
  • Bean, W.T., R. Stafford, and J.S. Brashares. 2011. The effects of small sample size and sample bias on threshold selection and accuracy assessment of species distribution models. Ecography 35:250-258.
  • Prugh, L., C. Stoner, C. Epps, W.T. Bean, W. Ripple, A. Laliberte, and J.S. Brashares. 2009. The rise of the mesopredator. Bioscience 59:779-791.
  • Wittemyer, G., P. Elsen, W.T. Bean, A.C. Burton, and J.S. Brashares. 2008. Accelerated human population growth at protected area edges. Science 321:123-126.
  • Bean, W.T., and E.W. Sanderson. 2007. Using a spatially explicit model to test scenarios of fire use by Native Americans: an example from the Harlem Plains, New York, NY. Ecological Modelling 211:301-308.